Specifically when they are level drained to the point that they "lose" wildshape? Or they "lose" the ability to wildshape into a huge creature (but were already a huge creature anyway)?
Energy Drain Relevant text:
The creature is also treated as one level lower for the purpose of level-dependent variables (such as spellcasting) for each negative level possessed.
prototype00
No? Fair enough. *phew*. So I get to whack the vampire in T-Rex mode then? Shiny.
prototype00
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It will shorten the duration you can stay in that form and if you were close to the end it might cause the Wild Shape to end prematurely as duration of the shape is a level dependent variable.
What you can turn into seems to be a level dependent variable so I might prevent from assuming the form of a huge creature if you no longer qualify but you would not lose the shape your in except as pointed out above.
I don't think there should be any effect on an already active Wild Shape, nor (as Jiggy pointed out) to whether you have the ability, or how many times per day you can use it. It would affect what forms you can use, and the spells you use to determine the effects of those forms, for any further uses of Wild Shape.
So if you were an 8th level Druid already in the form of a T-Rex, you're good if hit by a negative level - like any active spell on you. But if you were scouting as an eagle you won't be able to turn into one afterwards.
Edit: Revised based on reading. Since the ability functions as the spell, ongoing spell effects should not be affected. So nothing would happen with respect to an ongoing wild shape any more than a wizard's existing magnificent mansion spell would wink out if you level-drained the wizard enough.
I don't believe your ability to turn into more advanced forms would be affected. To echo Jiggy, "stuff that you can turn into" is based on level, but it isn't really a level-dependent variable in that sense.
Now, if the levels became permanent, then you would completely lose wildshape, at least as I understand it... but prior to the roll to see if it becomes permanent, you just get a -1 to all rolls basically.
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RPG Superstar 2012 Top 32
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Now, if the levels became permanent, then you would completely lose wildshape, at least as I understand it... but prior to the roll to see if it becomes permanent, you just get a -1 to all rolls basically.
I don't see anything in the rules suggesting that temporary and permanent negative levels are any different from each other in that regard.
RPG Superstar 2011 Top 32
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I don't think there should be any effect on an already active Wild Shape, nor (as Jiggy pointed out) to whether you have the ability, or how many times per day you can use it. It would affect what forms you can use, and the spells you use to determine the effects of those forms, for any further uses of Wild Shape.
So if you were an 8th level Druid already in the form of a T-Rex, you're good if hit by a negative level - like any active spell on you. But if you were scouting as an eagle you won't be able to turn into one afterwards.
This is how I see it, pretty much. It would have no effect on your already active Wild Shape, and would shorten the duration of future changes.
I can see arguments both ways about whether your available variety of forms is a level-dependent variable. I think I lean toward it not being one, inasmuch I don't want fighters to have to recalculate Weapon Training bonuses and clerics changing Channel Energy dice and so forth.
Now, if the levels became permanent, then you would completely lose wildshape, at least as I understand it... but prior to the roll to see if it becomes permanent, you just get a -1 to all rolls basically.
Some abilities and spells (such as raise dead) bestow permanent level drain on a creature. These are treated just like temporary negative levels, but they do not allow a new save each day to remove them.
Maybe you're getting this mixed up with the rules for permanent bonuses? I can rarely keep all of this straight.
In any case, you do lose some of your wild shape efficacy (namely duration & uses per day) whether the negative levels are temporary or permanent, but any currently active wild shape should be ineffective.