Can metamagic feats be applied to a cleric's domain spells? If so, does a higher level domain spell slot get used?
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The way that metamagic affects domain spells isn't much different from normal spells, but will change a little depending on whether that PC is a prepared spellcaster, a spontaneous caster, or using a metamagic rod or other item. Some feats may reduce or remove the cost of metamagic feats as well.
Most clerics are prepared casters and must apply the metamagic feat when preparing/recieving their spells. In this case, a higher-level domain slot will probably be needed.
I think they have the option of leaving the higher-level domain slot 'open' and unmemorized so that when the situation arrises, they can apply a metamagic feat to a lower-level memorized spell using the higher slot and a full round action.
Can metamagic feats be applied to a cleric's domain spells? If so, does a higher level domain spell slot get used?
Yes and yes. See the magic chapter about filling higher level spell slots with lower level spells.
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I think they have the option of leaving the higher-level domain slot 'open' and unmemorized so that when the situation arrises, they can apply a metamagic feat to a lower-level memorized spell using the higher slot and a full round action.
Please give me a source for that conversion thing, it would be very interesting to me. I know of no such rule, it seems like a house rule.
You can however, as always, leave a slot open and memorize a spell later.