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My first instinct with the Guinea Lion was to critique a predator's build on an herbivore. Predator builds are biologically expensive. There's a reason they eat meat. Also, lions can leap, so how do you expect to ranch them? You'd need huge fences, or a ceiling. This guinea lion idea is sooooo unrealistic.
Most likely by acclimating them to human contact and providing a steady food source. Cause you're right, they'd have to be voracious herbivores, which is what inspired the long distance movement penalty. They have to spend so much time eating they can't exist in large packs, nor can they travel far in a single day.
But remember, they aren't the result of evolution, but rather magical experimentation.
Also, I want a sharkhorse.